Arctic LTER Database

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Shaver, G. 1989. Above ground biomass in acidic tussock tundra experimental site, 1989, Arctic LTER, Toolik, Alaska. Arctic LTER, Marine Biological Lab, Woods Hole, Ma 02543. 1989gsttbm 

4 ) Acknowledgement. The Data User should acknowledge any institutional support or specific funding awards referenced in the metadata accompanying this dataset in any publications where the Data Set contributed significantly to its content. Acknowledgements should identify the supporting party, the party that received the support, and any identifying information such as grant numbers. For example:

Data sets were provided by the Arctic LTER. This material is based upon work supported by the National Science Foundation under Grants #DEB-981022, 9211775, 8702328; #OPP-9911278, 9911681, 9732281, 9615411, 9615563, 9615942, 9615949, 9400722, 9415411, 9318529; #BSR 9019055, 8806635, 8507493.

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Dataset URLs:METADATA: HTML, Rich Text, XML(EML compliant)
DATA: Comma Delimited, Excel file with Metadata and data, Dataset via LTER Data Portal
Dataset ID:1983_EVRT1980_TILLER_SIZE.02
Dataset Title:Tiller size measured on intact shoots in 1983 for the 1980-82 Eriophorum vaginatum reciprocal transplant experiment
Investigator 1: 
First Name:Ned
Last Name:Fetcher
Organization:Wilkes University
Address line 2:Institute for Environmental Science and Sustainabilty
Address line 3:84. W. South St.
Zip Code:18766
Investigator 2: 
First Name:James
Last Name:McGraw
Organization:West Virginia University
Address line 2:Department of Biology
Address line 3:53 Campus Drive
Zip Code:26506-6057
Investigator 3: 
First Name:Gaius
Last Name:Shaver
Organization:Marine Biological Laboratory
Address line 2:The Ecosystem Center
Address line 3:
City:Woods Hole
Zip Code:02543
Investigator 4: 
First Name:Cynthia
Last Name:Bennington
Organization:Stetson University
Address line 2:Department of Biology
Zip Code:32723
Associate Investigators:
Keywords:tiller size, Eriophorum vaginatum, tussock cottongrass, 1980 Eriophorum vaginatum transplant experiment, reciprocal transplant, populations
Abstract: These data were collected in August 1983 for tussocks transplanted in 1980-82 in a reciprocal transplant experiment and harvested in 2011. Important variables are garden name, source population, the number of green leaves, and the length of the longest leaf.
For questions about the Metadata and data contact the Investigators.
For information about this web site contact:
Arctic LTER Information Manager
The Ecosystems Center
Marine Biological Lab
7 MBL St
Woods Hole, MA 02543
Phone (508) 289 7496
Online URL:
Data File URL
Data File Name 1983_EVRT1980_TILLER_SIZE
Beginning Date 7/29/1983
End Date 8/24/1983
Number of Data Records 1539
Other Files to Reference
Availability Status 1
Quality Control Information
Maintenance Description
Log of Changes: Version 1: added to Arctic LTER database. Jim L 30Jan14
Version 2: Checked keywords against the LTER network preferred list and replaced non-preferred terms. Jim L 27Jan14
RESEARCH LOCATION:                        
Location Name Eagle Creek Transplant Garden No Name Creek Transplant Garden Coldfoot Transplant Garden Toolik Lake Transplant Garden Sagwon Transplant Garden Prudhoe Bay Transplant Garden Kuparik Transplant Garden Galbraith Transplant Garden Old Man Transplant Garden Atigun Camp Transplant Garden MS119 Transplant Garden  
Geographic Description Eagle Creek Tussock Tundra Site on hillside above Eagle Creek at Mile 101 of the Steece Highway| garden is about 500 m upslope from an old berm created by bulldozer. Coordinates are approximate. Coordinates and elevation for all sites are from Google Earth. The garden is in the middle of giant tussocks 100 m north and 165 m west of the bridge that crosses No Name Creek on the Dalton Highway. Access is easier by going along No Creek for 165 m and then heading north. The garden is 100 m north of the edge of the forest and 190 m east of the Dalton Highway. The garden is in the complex of "historic sites" at Toolik Lake. The garden is 50 m north of the upper parking area at the Sagwon Overlook pulloff. Site fomerly known as MS 127. The garden is 1.5 km west of the Kuparik Bridge and 420 m north of the road on the east bank of a stream. Note: Permission from BP and oil field safety training required to access this site. The garden is 120 m north of the Dalton Highway on the east side of the bridge. It consists of 2 rows of stakes. The garden is 60 m north of the road to the Galbraith Lake Airport. The garden is 470 m east of the Dalton Highway on the downhill side of the old pad for the Old Man construction camp. The Shaver Eriophorum vaginatum flower plots are located at the upper (south) end of Atigun Camp, just below a tributary stream that comes in form the west. To find them drive to just beyond the upper camp entrance and look up the hill. The flower plots are about 150 m up the hill. The transplanted tussocks are on the one side (north?) of the plots. The Shaver Eriophorum vaginatum flower plots are located just off access road APL119-2 (west side iof road). Walk down the access road about 50 m, then left over the tundra along a vehicle track about 200 m to the flower plots. The transplanted tussocks are between the flower plots and the haul road (closer to the flower plots).  
Location Bounding Box                        
West Bounding Coordinate                        
East Bounding Coordinate                        
North Bounding Coordinate                        
South Bounding Coordinate                        
OR if single point location                        
Latitude 65.433665 66.116744 67.258973 68.629538 69.424431 70.328353 68.648459 68.49629 66.460996 68.174583 68.760273  
Longitude -145.514261 -150.169792 -150.169153 -149.579858 -148.709647 -149.062722 -149.41033 -149.471807 -150.639054 -149.438122 -148.89143  
Elevation 762 167 319 759 269 8 735 831 470 1062 505  
Link to Google Map View on Google Map View on Google Map View on Google Map View on Google Map View on Google Map View on Google Map View on Google Map View on Google Map View on Google Map View on Google Map View on Google Map  
Organisms studied Eriophorum vaginatum
Methods:In 1980-1982, six transplant gardens were established along a latitudinal gradient in interior Alaska from Eagle Creek, AK, in the south to Prudhoe Bay, AK, in the north (Shaver et al. 1986) .Three sites, Toolik Lake (TL), Sagwon (SAG), and Prudhoe Bay (PB) are north of the continental divide and the remaining three, Eagle Creek (EC), No Name Creek (NN), and Coldfoot (CF), are south of the continental divide. Each garden consisted of 10 individual tussocks transplanted back to their home-site, as well as 10 individuals from each of the other transplant sites (n = 10; 6 populations x 6 sites x 10 replicates = 360 total individuals).

All gardens were censused in 1983. The number of flowers for each tussock was counted. For a haphazardly chosen subsample of tillers ( n = 3) on each tussock, the total number of green leaves was counted and the length of the longest leaf was measured from the top of the sheath. The width of the longest leaf was measured with an optical comparator. Old leaves that were initiated the previous year were identified by a dark band with dead tissue above it, whereas new leaves had no such band. A tiller size index was calculated by multiplying the length of the longest leaf by the total number of green leaves (usually 2, 3 or 4). This tiller index is highly correlated with tiller mass (Shaver, Fetcher & Chapin 1986).

In this census, we also measured tiller size index at five additional gardens, Kuparik River (KUP), Galbraith Lake (GAL), Old Man Camp (OLDMAN), Atigun Pass (ATIGUN), MS119 (MS119) and for two additional populations Kuparik (KUP) and Galbraith (GAL). In addition, we measured number of green leaves and length of the longest leaf for 10 untransplanted tussocks at each garden. These are designated by CONTROL.

Bennington CC, Fetcher N, Vavrek MC, Shaver GR, Cummings KJ, McGraw JB (2012) Home site advantage in two long-lived arctic plant species: results from two 30-year reciprocal transplant studies. Journal of Ecology 100:841-851

Shaver GR, Fetcher N, Chapin FS (1986) Growth and flowering in Eriophorum vaginatum - Annual and latitudinal variation. Ecology 67:1524-1535

Fetcher, N., and G. R. Shaver. 1990. Environmental sensitivity of ecotypes as a potential influence on primary productivity. American Naturalist 136:126-131.


Tiller size data are incorporated in Fetcher and Shaver 1990.
This data was collected with support from the School for Field Studies.

Data Table

Variable Name Variable Description Data Type Units DateTime Format Code Information Missing Value Code
Garden Name of the transplant garden text     EC= Eagle Creek| NN= No Name Creek| CF= Coldfoot| TL= Toolik Lake| SAG= Sagwon| PB= Prudhoe Bay| GAL= Galbraith| KUP= Kuparik| ATIGUN= Atigun Pass| OLDMAN= Old Man Camp| MS119= MS119 Site  
Source Name of the source population. text     EC= Eagle Creek| NN= No Name Creek| CF= Coldfoot| TL= Toolik Lake| SAG= Sagwon| PB= Prudhoe Bay| GAL= Galbraith| KUP= Kuparik| CONTROL= Local Untransplanted Tussock  
Tussock Number of tussock number dimensionless     ""=Missing or Not Measured
Flower Number of Inflorescences on the tussock number dimensionless     ""=Missing or Not Measured
IndexMean Mean of TillerIndex for the tussock number millimeter     ""=Missing or Not Measured
Comment The comments note the percentage of the tussock that had turned yellow/brown (senescent) and/or was grazed (presumably by lemmings or ptarmigan). text       ""=Missing or Not Measured
Sample Initials for the person who collected the data text     NF= Ned Fetcher| JBM= James B McGraw| CB= Cynthia Bennington| MV= Milan Vavrek| BB= Brian Bovard| APG= April Grossman ""=Missing or Not Measured
NumNewLeaves Number of New Green Leaves number dimensionless     ""=Missing or Not Measured
NumGreenLeaves Number of Old Green Leaves + Number of New Green Leaves number dimensionless     ""=Missing or Not Measured
LL Length of the longest leaf number millimeter     ""=Missing or Not Measured
WidthLL Width of the longest leaf number millimeter     ""=Missing or Not Measured
TillerIndex NumGreenLeaves * LL number millimeter     ""=Missing or Not Measured