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Primary production

Stream primary production

Throughout the summer, several indices of primary productivity are measured in control and enriched sections of each stream. One such parameter is the concentration of chlorophyll a in the epilithic algae, determined by periodically scrubbing streambed rocks at several stations and measuring chlorophyll levels in the scrubbate. Dr. Mike Miller is the principal investigator in charge of epilithic chlorophyll determination. Also, photosynthetic and respiratory rates of epilithic algae (both rivers) and bryophytes (Kuparuk River) are measured. The principal investigator in charge of these metabolic rates is Dr. Breck Bowden. Bryophytes, particularly mosses, have become increasingly important in the streams. Files containing photosynthetic and respiratory rates of mosses are also presented here. Dr. Breck Bowden is the principal investigator for the bryophyte studies.

The menu below provides a complete list of the files on epilithic chlorophyll biomass and primary producer metabolism. Consult the brief descriptor following each file name for the information contained in each file. Clicking on a file name will send you to a document file that describes the method in which the data were collected; from the document file, you can access the data file by clicking on the hypertext data file name.


  • 00pekchl - Sestonic and epilithic chlorophyll for the Kuparuk River during the summer of 2000.

  • 00peochl - Sestonic and epilithic chlorophyll for Oksrukuyik Creek during the summer of 2000.

  • 01pekchl - Sestonic and epilithic chlorophyll for the Kuparuk River during the summer of 2001.

  • 01peochl - Sestonic and epilithic chlorophyll for Oksrukuyik Creek during the summer of 2001.

  • 02pekchl - epilithic and sestonic chlorophyll for the Kuparuk River during the summer of 2002.

  • 02peochl - Epilithic and sestonic chlorophyll for Oksrukuyik Creek during the summer of 2002.

  • 03pekchl - epilithic (ugChl/cm^2) and sestonic (ug Chl/L) chlorophyll for the Kuparuk River during the summer of 2003.

  • 03peochl - Epilithic and sestonic chlorophyll for Oksrukuyik Creek during the summer of 2003.

  • 04pekchl - epilithic (ugChl/cm^2) and sestonic (ug Chl/L) chlorophyll a for the Kuparuk River during the summer of 2004.

  • 1983_2004pekchl - Epilithic chlorophyll for the Kuparuk River during the summers of 1983 - 2004.

  • 8395kschl - Sestonic chlorophyll for the Kuparuk River during the summers of 1983 - 1995.

  • 89bometa - Epilithic respiration, photosynthesis, chlorophyll a and nutrients from light/dark benthic metabolism chamber experiments. Samples were collected from stations within the CONTROL, +P, and +NP zones of the Kuparuk River during the +P and +NP experiments run during 1989. There are three groups of data in this file. The first group was collected by Jane Tucker, under the supervision of Bruce Peterson, in early July. Measurements were only done on riffle rocks. No chlorophyll a or nutrient data exist for these runs. The second set was collected in late July by Bowden and Finlay. The second group was collected in early August by Finlay. No nutrient data are available for these runs.

  • 9095oechl - Epilithic chlorophyll for Oks Creek during the summers of 1990 - 1995.

  • 9095oschl - Sestonic chlorophyll for Oks Creek during the summers of 1990 - 1995.

  • 90bometa - Epilithic respiration, photosynthesis, and chlorophyll a from benthic metabolism chamber measurements. Samples were collected from pools and riffles within the CONTROL and Phosphorus (P) enriched zones of the Kuparuk River and similar zones of Oksrukuyik Creek. There was no P added to Oksrukuyik Creek in 1990; data from this year will be used as a pre-enrichment, "baseline" comparison of unmanipulated control and "enriched" reaches.

  • 91bometa - Epilithic respiration, photosynthesis, and chlorophyll a from benthic metabolism measurements. Samples were collected from pools and riffles within the CONTROL and Phosphorus- (P) enriched zones of the Kuparuk River. Samples were collected from pools, runs, and riffles in similar zones of Oksrukuyik Creek. Nitrogen was not added to either river this year. 1991 was the first year for P fertilization of Oksrukuyik Creek. Benthic metabolism and chlorophyll a measurements made in this stream in 1990 may be used to compare the unfertilized "fertilized" zone and the control zone. These data are contained in the following files: "90BOMETA.DOC" AND "90BOMETA.DAT".

  • 92bometa - Epilithic respiration, photosynthesis, and chlorophyll_a from benthic metabolism measurements. Samples were collected from pools and riffles within the CONTROL and PHOSPHORUS (P) enriched zones of the Kuparuk River and from similar zones of Oksrukuyik Creek. There were no nitrogen additions to either river this year. P fertilization began on 25 June this year, several days earlier than usual. 1992 was the SECOND year for P fertilization of Oksrukuyik Creek. Benthic metabolism and chlorophyll_a measurements in this stream in 1990 may be used to compare the unfertilized "fertilized" zone and the control zone. These data are contained in the following files: "90BOMETA.DOC" and "90BOMETA.DAT". First-year fertilization response data for Oksrukuyik Creek may be found in files "91BOMETA.DOC" and "91BOMETA.DAT".

  • 92bomocb - Cover biomass measurements used to estimate bryophyte biomass in the Kuparuk River. Hygrohypnum and Schistidium were the only bryophytes harvested for this measurement from the Kuparuk River's fertilized reach. Harvests were done in August

  • 92bomogr - Bryophyte growth data from the Kuparuk River. We measured growth of Hygrohypnum spp. and Fontinalis neomexicana clumps and stem tips in control and fertilized (PO4 phosphorus), pools and riffles.

  • 92bomont - Bryophyte (Hygrohypnum, Schistidium, and Fontinalis) tissue phosphorus and nitrogen concentrations in control and phosphorus fertilized reaches of the Kuparuk River in 1992.

  • 92bomopt - Point transects used to estimate epilithic bryophyte and macroalgal cover of the Kuparuk River. Hygrohypnum, Fontinalis, and Schistidium were the dominant bryophytes encountered. Transects were done July (early) and July and August (late) in both fertilized (phosphorus-enriched) and control riffles of the Kuparuk.

  • 93bometa - Epilithic respiration, photosynthesis, and chlorophyll a from benthic metabolism measurements. Samples were collected from pools and riffles within the control and enriched zones of the Kuparuk River and from similar zones of Oksrukuyik Creek. Phosphorus (P) was the only nutrient added to the Kuparuk River this year but nitrogen (N) and P were added at one site to Oksrukuyik Creek. Fertilization began on 25 June this year. 1993 was the third year for P fertilization of Oksrukuyik Creek. Benthic metabolism and chlorophyll a measurements in this stream in 1990 may be used to compare the unfertilized "fertilized" zone and the control zone. These data are contained in the following files: "90BOMETA.DOC" and "90BOMETA.DAT". First- and second-year fertilization response data for Oksrukuyik Creek may be found in files "91BOMETA.DOC", "91BOMETA.DAT", "92BOMETA.DOC" and "92BOMETA.DAT".

  • 93bomogr - Bryophyte growth data from the Kuparuk River. We measured growth of Hygrohypnum spp. clumps in control and fertilized (PO4 phosphorus), pools and riffles.

  • 93bomopt - Point transects used to estimate epilithic bryophyte and macroalgal cover of the Kuparuk River and Oksrukuyik Creek. Hygrohypnum spp., Fontinalis neomexicana, and Schistidium agassizii were the dominant bryophytes encountered in the Kuparuk River; bryophyte cover was very low in Oksrukuyik Creek, predominately composed of Solenostoma spp. Transects were done June and July (early) and July and August (late) in both fertilized and control riffles of the Kuparuk River and Oksrukuyik Creek. Pool and run environments in both streams were sampled less intensively. Riffles upstream (-15 to -9 km) and downstream (+6 to +20 km) of the Kuparuk experimental reach were sampled in late July.

  • 94bometa - Epilithic respiration, photosynthesis, and chlorophyll a from benthic metabolism measurements. Samples were collected from pools and riffles within the control and enriched zones of the Kuparuk River and from similar zones of Oksrukuyik Creek. Samples were collected from riffles of the 1994 "New Reach" (Blueberry Creek). Phosphorus (P) was the only nutrient added to the Kuparuk River this year but nitrogen (N) and P were added at one site to Oksrukuyik Creek. Fertilization began on 25 June this year. 1994 was the fourth year for P fertilization of Oksrukuyik Creek. Benthic metabolism and chlorophyll a measurements in this stream in 1990 may be used to compare the unfertilized "fertilized" zone and the control zone. These data are contained in the following files: "90BOMETA.DOC" and "90BOMETA.DAT".

  • 94bomopt - Point transects used to estimate epilithic bryophyte and macroalgal cover of the Kuparuk River, Oksrukuyik Creek, and the 1994 new study reach, Blueberry Creek. Hygrohypnum spp., Fontinalis neomexicana, and Schistidium agassizii were the dominant bryophytes encountered in the Kuparuk River; bryophyte cover was very low in Oksrukuyik Creek and Blueberry Creek, predominately composed of Solenostoma spp. and Schistidium agassizii, respectively. Transects were done in only in July in both fertilized and reference riffles of the Kuparuk River; in June and July in reference and fertilized riffles of Oksrukuyik Creek; and once in July throughout the new study reach of Blueberry Creek.

  • 94mibchl - Sestonic and epilithic chlorophyll for Blueberry Cr. (I8 outlet) during the summer of 1994.

  • 95bometa - Bryophyte and epilithic algal respiration, photosynthesis versus irradiance, and total chlorophyll from Kuparuk River and Oksrukuyik Creek benthic metabolism measurements.

  • 95bomocb - Cover biomass measurements used to estimate bryophyte biomass in the Kuparuk River. Hygrohypnum and Schistidium were the only bryophytes harvested for this measurement from the Kuparuk River's fertilized reach. Harvests were done in August.

  • 95bomont - Bryophyte tissue phosphorus and nitrogen concentrations in phosphorus fertilized reaches of the Kuparuk River in 1995.

  • 95bomopt - Point transects used to estimate epilithic bryophyte and macroalgal cover of the Kuparuk River and Oksrukuyik Creek. Hygrohypnum, Fontinalis, and Schistidium were the dominant bryophytes encountered in the Kuparuk River's fertilized reach. Transects were done June and August in both fertilized (phosphorus-enriched) and reference reaches of the Kuparuk and once in July in both fertilized (phosphorus-enriched) and reference reaches of Oksrukuyik Creek.

  • 96bometa - Respiration, photosynthesis versus irradiance and temperature, and total chlorophyll from metabolism measurements of bryophytes collected from the Kuparuk River. Metabolic measurements of epilithic algae at constant light and temperature for the Kuparuk River and Oksrukuyik Creek are included. Further, in 1996 we measured epilithic algal metabolism in various habitats at the Mouth of the Kuparuk River. These measurements are include in this data file.

  • 96bomocb - Cover biomass measurements used to estimate Hygrohypnum biomass in the Kuparuk River. Hygrohypnum was the only bryophyte harvested for this measurement from the Kuparuk River's fertilized and recovery reaches. Harvests were done in June and August

  • 96bomoct - Chlorophyll concentrations of Schistidium agassizii and Hygrohypnum spp. retrieved from the Kuparuk River. In 1996, weekly, from 22-Jun to 10-Aug, collections of bryophytes from 3 riffles each treatment reach (i.e. reference, recovery, and fertilized).

  • 96bomopt - Point transects used to estimate bryophyte and macroalgal cover of the Kuparuk River and Oksrukuyik Creek. Hygrohypnum, Fontinalis, and Schistidium were the dominant bryophytes encountered in the Kuparuk River's fertilized reach. Transects were done in June and August in both fertilized (phosphorus-enriched), recovery, and reference reaches of the Kuparuk. Transects were done in late June and late July in both fertilized (phosphorus-enriched) and reference reaches of Oksrukuyik Creek.

  • 96mikchl - Sestonic and epilithic chlorophyll for the Kuparuk River during the summer of 1996.

  • 96miochl - Sestonic and epilithic chlorophyll for Oksrukuyik Creek during the summer of 1996.

  • 97pekchl - Sestonic and epilithic chlorophyll for the Kuparuk River during the summer of 1997.

  • 97peochl - Sestonic and epilithic chlorophyll for Oksrukuyik Creek during the summer of 1997.

  • 98pekchl - Sestonic and epilithic chlorophyll for the Kuparuk River during the summer of 1998.

  • 98peochl - Sestonic and epilithic chlorophyll for Oksrukuyik Creek during the summer of 1998.

  • 99pekchl - Sestonic and epilithic chlorophyll for the Kuparuk River during the summer of 1999.

  • 99peochl - Sestonic and epilithic chlorophyll for Oksrukuyik Creek during the summer of 1999.

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